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Underlined = undergraduate co-authors; asterisks = equal contribution

Link to Google Scholar page here

de Pablo, L.X., Carleton, A., Modarres-Sadeghi, Y., and Clotfelter, E.D. In press. Substrate interactions and free-swimming dynamics in the crayfish escape response. Integrative Organismal Biology. 

Rothberg, S.E., Wolf, S.E. and Clotfelter, E.D. 2024. Social network connections are positively related to temperature in winter flocks of black-capped chickadees. Animal Behaviour 210: 213-224.

Wolf, S.E., Woodruff, M.J., Chang van Oordt, D.A., Clotfelter, E.D., Cristol, D.A., Derryberry, E.P., Ferguson, S.M., Stanback, M.A., Taff, C.C., Vitousek, M.N., Westneat, D.F. and Rosvall, K.A. 2024. Among-population variation in telomere regulatory proteins and their potential role as hidden drivers of intraspecific variation in life history. Journal of Animal Ecology.

Benson, D.M. and Clotfelter, E.D. 2023. Size, but not sex, predicts pinch force and exoskeleton mechanical properties in crayfish of the genus Faxonius. Invertebrate Biology.

Dunn, P.O., Ahmed, I., Armstrong, E., Barlow, N., Barnard, M.A., Belisle, M., Benson, T.J., Berzins, L.L., Boynton, C.K., Anders Brown, T., Cady, M., Cameron, K., Chen, X., Clark, R.G., Clotfelter, E.D., et al. 2023. Extensive regional variation in the phenology of insects and their response to temperature across North America. Ecology 104: e4036.

Wolf, S.E., Zhang, S. and Clotfelter, E.D. 2023. Experimental ectoparasite removal has a sex-specific effect on nestling telomere length. Ecology and Evolution. doi: 10.1002/ece3.9861.

Earley, R.L., Anderson, C.T., Moscicki, M.K., Norton, B.B., Brown, A.C. and Clotfelter, E.D. 2020. Carotenoid availability and tradeoffs in female convict cichlids, a reverse sexually-dichromatic fish. Environmental Biology of Fishes 103: 1541-1552.

Elliott, H., Smith, C. and Clotfelter, E.D. 2020. Hummingbird frugivory in a Costa Rican cloud forest. Ornitología Neotropical 31: 71-75.


Hunyadi, J., Currier, T., Modarres-Sadeghi, Y., Flammang, B.E. and Clotfelter, E.D. 2020. Morphology, performance and fluid dynamics of the crayfish escape response. Journal of Experimental Biology. doi:10.1242/jeb.219873. 

* Article featured in “Inside JEB.” doi: 10.1242/jeb.231969


Sun, N.W., Goodwin, S.E., Griego, M.S., Gerson, A.R. and Clotfelter, E.D. 2019. Does blood loss explain higher resting metabolic rates in nestling birds with hematophagous ectoparasites? Journal of Avian Biology.

DeSimone*, J.G., Clotfelter*, E.D., Black, E.C. and Knutie, S.A. 2018. Avoidance, tolerance, and resistance to ectoparasites in nestling and adult tree swallows. Journal of Avian Biology 49: jav-01641.

Malavé, B.M., Styga, J.M. and Clotfelter, E.D. 2018. Size, shape, and sex-dependent variation in force production by crayfish chelae. Journal of Morphology 279: 312-318.

Clotfelter, E.D., Johnson, E.M., Anderson, C. and Earley, R.L. 2017. Evidence for male alternative reproductive tactics in convict cichlids (Amatitlania siquia) in Lake Xiloá, Nicaragua. Environmental Biology of Fishes 100: 655-662.

Brown, A.C., Cahn, M.D., Choi, S. and Clotfelter, E.D. 2016. Dietary carotenoids and bacterial infection in wild and domestic convict cichlids (Amatitlania spp.). Environmental Biology of Fishes 99: 439-449. 10.1007/s10641-016-0485-x

Anderson, C., Jones, R., Moscicki, M., Clotfelter, E.D., and Earley, R.L. 2016. Seeing orange: breeding convict cichlids exhibit heightened aggression against more colorful intruders. Behavioral Ecology and Sociobiology 70: 647-657. 10.1007/s00265-016-2085-3


Cahn, M.D., Brown, A.C. and Clotfelter, E.D. 2015. Guanine-based structural coloration as an indicator of oxidative stress in a cichlid fish. Journal of Experimental Zoology A 323: 359-367.

Sullivan, M., Brown, A.C. and Clotfelter, E.D. 2014. Dietary carotenoids do not improve motility or antioxidant capacity in cichlid fish sperm.  Fish Physiology and Biochemistry 40: 1399-1405.

Sefc, K., Brown, A.C. and Clotfelter, E.D. 2014. Carotenoid-based coloration in cichlid fishes. Comparative Biochemistry and Physiology A 173: 42-51.


Montgomery, T.M., Brown, A.C., Gendelman, H.K., Ota, M. and Clotfelter, E.D. 2014. Exposure to 17α-ethinyl estradiol decreases motility and ATP in sperm of male fighting fish Betta splendens. Environmental Toxicology 29: 243-252.

Brown, A.C., Stevenson, L.M., Leonard, H.M., Nieves-Puigdoller, K., Clotfelter, E.D. 2014. Phytoestrogens β-sitosterol and genistein have limited effects on reproductive endpoints in a female fish, Betta splendens. BioMed Research International: Toxicology doi:/10.1155/2014/681396


Clotfelter, E.D. and Gendelman, H. 2014. Exposure to environmentally relevant concentrations of genistein during activation does not affect sperm motility in the fighting fish Betta splendens. BioMed Research International: Toxicology. doi: 10.1155/2014/865741


Brown, A.C., Leonard, H.M., McGraw, K.J. and Clotfelter, E.D. 2014. Maternal effects of carotenoid supplementation in an ornamented cichlid fish (Amantitlania siquia). Functional Ecology 28: 612-620.


Clotfelter, E.D., Lapidus, S.J.H. and Brown, A.C. 2013. The effects of temperature and dissolved oxygen on antioxidant defences and oxidative damage in the fathead minnow Pimephales promelas.  Journal of Fish Biology 82: 1086-1092.


Brown, A.C., McGraw, K.J. and Clotfelter, E.D. 2013. Dietary carotenoids increase yellow non-pigment coloration of female convict cichlids (Amantitlania nigrofasciata). Physiological and Biochemical Zoology 86: 312-322.


Brown, A.C. and Clotfelter, E.D. 2012. Fighting fish (Betta splendens) bubble nests do not inhibit microbial growth. Journal of Experimental Zoology 317: 481-487. (Cover story)


Stevenson, L.M., Brown, A.C., Montgomery, T.M. and Clotfelter, E.D. 2011.  Reproductive consequences of exposure to waterborne phytoestrogens in male fighting fish Betta splendens. Archives of Environmental Contamination and Toxicology 60: 501-510.


Clotfelter, E.D., McNitt, M.M., Carpenter, R.E. and Summers, C. 2010. Modulation of monoamine neurotransmitters in fighting fish Betta splendens exposed to waterborne phytoestrogens. Fish Physiology and Biochemistry 36: 933-943.


Ardia, D.R., Pérez, J.H. and Clotfelter, E.D. 2010. Experimental cooling during incubation leads to reduced innate immunity and body condition in nestling tree swallows.  Proceedings of the Royal Society of London B 277: 1881-1888.


Lin, S.M., Nieves-Puigdoller, K., Brown, A.C., McGraw, K.J. and Clotfelter, E.D. 2010. Testing the carotenoid trade-off hypothesis in the polychromatic Midas cichlid, Amphilophus citrinellus. Physiological and Biochemical Zoology 83: 333-342.


Morrison, E.S., Ardia, D.R. and Clotfelter, E.D. 2009. Cross-fostering reveals sources of variation in innate immunity and hematocrit in nestling tree swallows. Journal of Avian Biology 40: 573-578.


Kuperberg, E.S., Brown, A.C. and Clotfelter, E.D. 2009. Body condition in male Betta splendens does not predict their ability to perform opercular displays under hypoxic conditions. Ethology 115: 1182-1189.


Blakeslee, C., McRobert, S.P., Brown, A.C. and Clotfelter, E.D. 2009. The effect of body coloration and group size on social partner preferences in female fighting fish (Betta splendens). Behavioural Processes 80: 157-161.


Ardia, D.R., Pérez, J.H., Chad, E.K. and Clotfelter, E.D. 2009. Temperature and life history: experimental heating leads female tree swallows to modulate egg temperature and incubation behavior.  Journal of Animal Ecology 78: 3-14.


Pérez, J.H., Ardia, D.R., Chad, E.K. and Clotfelter, E.D. 2008. Experimental heating reveals nest temperature affects nestling condition in tree swallows (Tachycineta bicolor). Biology Letters 4: 468-471.


Clotfelter, E.D. 2008. Frequency-dependent performance and handedness in professional baseball players. Journal of Comparative Psychology 122: 68-72.


Clotfelter, E.D. and Hollis, K.L. 2008. Cognition in domestic dogs: object permanence and social cueing. The American Biology Teacher 70: 293-298.


Clotfelter*, E.D., Ardia*, D.R. and McGraw, K.J. 2007. Red fish, blue fish: trade-offs between pigmentation and immunity in Betta splendens. Behavioral Ecology 18: 1139-1145.


Clotfelter*, E.D., Pedersen*, A.B., Cranford, J.A., Ram, N., Snajdr, E.A., Nolan Jr., V. and Ketterson, E.D. 2007. Acorn mast drives long-term dynamics of rodent and songbird populations. Oecologia 154: 493-503.


Clotfelter, E.D., O’Hare, E.P., McNitt, M.M., Carpenter, R.E. and Summers, C.H. 2007. Serotonin decreases aggression via 5-HT1A receptors in the fighting fish Betta splendens. Pharmacology, Biochemistry and Behavior 87: 222-231.


Ardia, D.R. and Clotfelter, E.D. 2007. Individual quality and age affect responses to an energetic constraint in a cavity-nesting bird. Behavioral Ecology 18: 259-266.


Clotfelter, E.D., Chandler, C.R., Nolan, V. Jr. and Ketterson, E.D. 2007. The influence of exogenous testosterone on the dynamics of nestling provisioning in dark-eyed juncos. Ethology 113: 18-25.


Clotfelter, E.D. and Kuperberg, E.S. 2007. Cerebral lateralization and its relationship to phylogeny and aggression in anabantoid fishes. Brain, Behavior and Evolution 69: 169-175.


Clotfelter, E.D., Curren, L.J. and Murphy, C.E. 2006. Mate choice and spawning success in the fighting fish Betta splendens: the importance of body size, display behavior and nest size. Ethology 112: 1170-1178.


Ardia, D.R., Pérez, J.H. and Clotfelter, E.D. 2006. Nest box orientation affects internal temperature and nest site selection by tree swallows. Journal of Field Ornithology 77: 339-344.


Clotfelter, E.D. and Rodriguez, A.C. 2006. Behavioral changes in fish exposed to phytoestrogens. Environmental Pollution 144: 833-839.


Ardia, D.R. and Clotfelter, E.D. 2006. The novel application of an immunological technique reveals the immunosuppressive effect of phytoestrogens in Betta splendens. Journal of Fish Biology 68 (Suppl. A): 144-149.


Snekser, J.L., McRobert, S.P. and Clotfelter, E.D. 2006. Social partner preferences of male and female fighting fish (Betta splendens). Behavioural Processes 72: 38-41.


Snekser, J.L., McRobert, S.P., Murphy, C.E. and Clotfelter, E.D. 2006. Aggregation behavior in wildtype and transgenic zebrafish. Ethology 112: 181-187.


Sampieri, S.J. and Clotfelter, E.D. 2005. Temporal and geographic patterns of age-specific gull plumage in relation to potential exposure to polychlorinated biphenyls. Auk 122: 915-921.


Clotfelter, E.D., Bell, A.M. and LeVering, K. 2004. The role of animal behaviour in the study of endocrine-disrupting chemicals. Animal Behaviour 68: 465-476.


Clotfelter, E.D., O’Neal, D.M., Gaudioso, J.M., Casto, J.M., Parker-Renga, I.M., Snajdr, E.A., Duffy, D.L., Nolan, V. Jr. and Ketterson, E.D. 2004. Consequences of elevating plasma testosterone in females of a socially monogamous songbird: evidence of constraints on male evolution? Hormones and Behavior 46: 171-178.


Ingham, R.R., Gesualdi, D.A., Toth, C.R. and Clotfelter, E.D. 2004. Effects of genistein on growth and development of aquatic vertebrates. Bulletin of Environmental Contamination and Toxicology 72: 625-631.


Clotfelter, E.D. and Paolino, A.D. 2003. Bystanders to contests between conspecifics are primed for increased aggression in male fighting fish. Animal Behaviour 66: 343-347.


Whittingham, L.A., Dunn, P.O. and Clotfelter, E.D. 2003. Parental allocation of food to nestling tree swallows: the influence of nestling behaviour, sex and paternity. Animal Behaviour 65: 1203-1210.


Clotfelter, E.D., Schubert, K.A., Nolan, V., Jr. and Ketterson, E.D. 2003. Mouth color signals thermal state of nestling dark-eyed juncos (Junco hyemalis). Ethology 109: 171-182.


Clotfelter, E.D., Nolan, V., Jr. and Ketterson, E.D. 2001. The effects of experimentally elevated testosterone and food deprivation on food consumption and prey size preferences in male dark-eyed juncos (Junco hyemalis, Emberizidae: Passeriformes). Ethology 107: 439-449.  


Clotfelter, E.D., Whittingham, L.A. and Dunn, P.O. 2000. Laying order, hatching asynchrony and nestling body mass in tree swallows Tachycineta bicolor. Journal of Avian Biology 31: 329-334.


Clotfelter, E.D. and Yasukawa, K. 1999. The effect of aggregated nesting on red-winged blackbird nest success and brood parasitism by brown-headed cowbirds. Condor 101: 729-736.


Clotfelter, E.D. and Yasukawa, K. 1999. Impact of brood parasitism by brown-headed cowbirds on red-winged blackbird reproductive success. Condor 101: 105-114.


Clotfelter, E.D. and Yasukawa, K. 1999. The function of early onset of nocturnal incubation in red‑winged blackbirds. Auk 116: 417-426.


Clotfelter, E.D., Yasukawa, K. and Newsome, R.D. 1999. The effects of prescribed burning and habitat edges on brown-headed cowbird parasitism of red-winged blackbirds. Studies in Avian Biology 18: 275-281.


Clotfelter, E.D. 1998. What cues do brown-headed cowbirds use to locate red-winged blackbird host nests? Animal Behaviour 55: 1181-1189.


Clotfelter, E.D. 1997. Red‑winged blackbird parental investment following brood parasitism by brown‑headed cowbirds: is parentage important? Behavioral Ecology and Sociobiology 41: 193-201.


Clotfelter, E.D. 1996. Mechanisms of facultative sex ratio variation in zebra finches (Taeniopygia guttata). Auk 113: 441‑449.


Clotfelter, E.D. 1995. Courtship displaying and intrasexual competition in the bronzed cowbird. Condor 97: 816‑818.


Clotfelter, E.D. and Brush, T. 1995. Unusual parasitism by the bronzed cowbird. Condor 97: 814‑815.

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